Wuerhosaurus is a genus of stegosaurid dinosaur from the Early Cretaceous Period of China and Mongolia. As such, it was one of the last genera of stegosaurians known to have existed, since most others lived in the late Jurassic.
Discovery and species[]
Wuerhosaurus homheni is the type species, described by Dong Zhiming in 1973 from the Tugulu Group in Xinjiang, western China. The generic name is derived from the city of Wuerho. Three separate localities in the Wuerho Valley were discovered to contain material from the new stegosaur: 64043-5, 64043 and 64045. The remains consisted of the holotype, Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) V.4006, a skull-less fragmentary skeleton, and the paratype IVPP V.4007. Holotype material includes a mostly complete pelvis and sacrum lacking the ischium, the first caudal vertebrae, two dorsal vertebrae, a scapulocoracoid, humerus and phalanx, as well as two dermal plates. Three posterior caudal vertebrae from the tail and a partial ulna of a second individual form the paratype, and Dong referred a partial ischium from a third locality to Wuerhosaurus.
A smaller stegosaur from the Ejinhoro Formation in the Ordos Basin in Inner Mongolia was found in 1988. When the specimen (IVPP V.6877) was described by Dong in 1993, it was named W. ordosensis, as it was from a similar age and had a similar anatomy. The holotype of the species includes a nearly complete torso, consisting of three cervical vertebrae, all eleven dorsal vertebrae (with attached ribs), a complete sacrum with a right ilium, and the first five caudal vertebrae, all articulated. An additional dorsal vertebra and dermal plate were referred to the taxon when it was named. In 2014 Ulansky named a new species of Wuerhosaurus, "W. mongoliensis" for vertebrae and pelvic material, but the name is an invalid nomen nudum. It was formally described as Mongolostegus in 2018.
Description[]
Wuerhosaurus homheni was probably a broad-bodied animal. Gregory S. Paul in 2016 estimated the length at 7 m (23 ft) and the weight at 4 tonnes (8,800 lb). Only a few scattered bones have been found, making a full restoration difficult. Its dorsal plates were at first thought to have been much rounder or flatter than other stegosaurids, but Maidment established this was an illusion caused by breakage: their actual form is unknown. W. homheni had a pelvis of which the front of the ilia strongly flared outwards indicating a very broad belly. The neural spines on the tail base were exceptionally tall.
W. ordosensis was estimated by Paul to have been 5 m (16 ft) long and weigh 1.2 tonnes (2,600 lb). It too has a broad pelvis but the neural spines are shorter. The neck seems to have been relatively long.
Classification[]
Phylogeny[]
Wuerhosaurus is one of the most derived stegosaurians, being closely related to either Dacentrurus and Hesperosaurus, or Hesperosaurus and Stegosaurus, depending on phylogenetic analysis. Carpenter et al. (2001) recovered Wuerhosaurus in the former relationship, close to Hesperosaurus and Dacentrurus as basal in Stegosauridae. Wuerhosaurus was recovered in a different position by Escaso et al. (2007), still related to Hesperosaurus, but basal to a clade of Lexovisaurus and Stegosaurus. Maidment et al. (2008) recovered a different placement with Wuerhosaurus as being in a clade of taxa in derived Stegosaurinae, most closely related to Hesperosaurus and then Stegosaurus. More recently, Maidment (2017) elaborated upon her earlier analyses, and instead resolved Wuerhosaurus as closest to Stegosaurus, with Hesperosaurus being more closely related to Miragaia. These results are shown below.
| Stegosauria |
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Synonymy with Stegosaurus[]
Maidment and colleagues proposed in 2008 that Wuerhosaurus was a junior synonym of Stegosaurus, with W. homheni being renamed Stegosaurus homheni, and W. ordosensis being a dubious taxon. W. homheni was synonymized because the holotype that could be located was similar to Stegosaurus, and because of its phylogenetic position. Wuerhosaurus placed between Stegosaurus and Hesperosaurus, the latter being considered a species of Stegosaurus because of its age, location, and anatomy. Since Wuerhosaurus placed between two possible Stegosaurus species, Maidment et al. synonymized the taxon as well. Wuerhosaurus ordosensis was considered to be a nomen dubium because the holotype could not be found in the IVPP collections. The original description did not mention any valid diagnostic traits, and no other description provided features either, so Maidment et al. considered the taxon undiagnostic.[10] This opinion has been contested, however, by Carpenter (2010[12]). He discussed how the diagnoses and features used by Maidment et al. were inconsistent and generalized, with Wuerhosaurus homheni bearing numerous differences. As such, Carpenter advocated for the separation of both Hesperosaurus and Wuerhosaurus from Stegosaurus, and the separation of the different Stegosaurus species from S. armatus.