This list of pterosaurs is a comprehensive listing of all genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (nomen dubium), or were not formally published (nomen nudum), as well as junior synonyms of more established names, and genera that are no longer considered pterosaurian. The list currently includes 157 genera.
[edit] Scope and terminology
Edit
There is no official, canonical list of pterosaur genera, but the most thorough attempts can be found at the Pterosauria section of Mikko Haaramo's Phylogeny Archive,[1] the Genus Index at Mike Hanson's The Pterosauria,[2] supplemented by the Pterosaur Species List,[3] and in the fourth supplement of Donald F. Glut's Dinosaurs: The Encyclopedia series.[4]
[edit] Authors and Year
Edit
The authors column lists the authors of the formal description responsible for the erection of the genus listed. They are not necessarily the same as the authors of the type species as sometimes a species from one genus is determined sufficiently distinct to warrant the erection of a new genus to house it. If this is the case, only the latter authors will be listed. The year column notes the year the description was published.
Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include:
- Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior synonyms are generally used, except by special decision of the ICZN, but junior synonyms cannot be used again, even if deprecated. Junior synonymy is often subjective, unless the genera described were both based on the same type specimen.
- Nomen nudum (Latin for "naked name"): A name that has appeared in print but has not yet been formally published by the standards of the ICZN. Nomina nuda (the plural form) are invalid, and are therefore not italicized as a proper generic name would be. If the name is later formally published, that name is no longer a nomen nudum and will be italicized on this list. Often, the formally published name will differ from any nomina nuda that describe the same specimen.
- Preoccupied name: A name that is formally published, but which has already been used for another taxon. This second use is invalid (as are all subsequent uses) and the name must be replaced. As preoccupied names are not valid generic names, they will also go unitalicized on this list.
- Nomen dubium (Latin for "dubious name"): A name describing a fossil with no unique diagnostic features. As this can be an extremely subjective and controversial designation, this term is not used on this list.
The age column denotes the epoch of geologic time to which the fossils date. The abbreviations are as follows.
Genera that are invalid, misidentified, or otherwise do not represent a valid pterosaur are listed as age N/A because there was never a time in which a pterosaur by that generic name actually lived.
[edit] Location and Notes
Edit
The location column designates the geographic region where remains of the relevant genus have been found. The regions used are continents except in the case of smaller landmasses (eg. Cuba). Political bodies, being non-existent in the Mesozoic are not used to indicate genera locations. Genera that are invalid, misidentified, or otherwise do not represent a valid pterosaur are listed as location N/A because there was never a place in which a pterosaur by that generic name actually lived. The notes column is a collection of annotations on the scientific significance and taxonomic history of listed genera, as well as elaborations on the information presented in other columns.
|
|
|
|
| Genus[1]
| Authors[2]
| Year[3]
| Status[4]
| Age[5]
| Location[6]
| Notes
|
|
Aetodactylus[5]
|
Myers
|
2010
| Valid.
|
8 Late Cretaceous
|
N. America
|
|
|
Aidachar[6]
|
Nesov
|
1981
| Misidentification.
|
N/A
|
N/A
|
Actually a teleost fish whose remains were originally mistaken for jaw fragments of a ctenochasmatid. The mistake was corrected in 1986.
|
|
Amblydectes
|
Hooley
|
1870
|
Jr. synonym.
|
N/A
|
N/A
|
Junior synonym of Coloborhynchus.
|
|
Angustinaripterus[7]
|
He
Xinlu
et al.
|
1983
| Valid.
|
5 Middle Jurassic
|
Asia
|
Known from a single skull recovered from the Dashanpu Formation. Its Dorygnathus-like teeth suggest it had a piscivorous diet.
|
|
Anhanguera[8]
|
Campos
Kellner
|
1985
| Valid.
|
7 Early Cretaceous
|
S. America
|
A piscivorous ornithocheirid from the Santana Formation. A study of its anatomy helped resolve controversy regarding pterodacyloids' posture while on the ground.
|
|
Anurognathus[9]
|
Döderlein
|
1923
| Valid.
|
6 Late Jurassic.[10]
|
Europe.[10]
|
A tiny (50 cm wingspan) insectivore known only from two skeletons recovered from the Solnhofen Formation.
|
|
Apatomerus[11]
|
Williston
|
1903
|
Misidentification.[2]
|
7 Lower Cretaceous
|
N. America
|
Probably a misidentified plesiosaur.[2]
|
|
Arambourgiania[12]
|
Nesov
Kanznyshkina
Cherepanov
|
1987
| Valid.
|
8 Late Cretaceous
|
Asia
|
|
|
Aralazhdarcho[13]
|
Averianov
|
2007
| Valid.
|
8 Late Cretaceous
|
Asia
|
|
|
Araripedactylus
|
Wellnhofer
|
1977
| Valid.
|
7 Early Cretaceous
|
S. America
|
|
|
Araripesaurus
|
Price
|
1971
| Valid.
|
7 Early Cretaceous
|
S. America
|
|
|
Arthurdactylus
|
Frey
Martill
|
1994
| Valid.
|
7 Early Cretaceous
|
S. America
|
|
|
Austriadactylus[7]
|
Dalla Vecchia
Wild
Reitner
|
2002
| Valid.
|
3 Upper Triassic
|
Europe
|
|
|
Avgodectes[14]
|
Peters
|
2004
|
Jr. synonym.
|
N/A
|
N/A
|
Probable junior synonym of Haopterus.
|
|
Azhdarcho[15]
|
Nesov
|
1984
| Valid.
|
8 Late Cretaceous
|
Asia
|
|
|
Bakonydraco[16]
|
Weishampel
Jianu
|
2005
|
Valid.
|
8 Late Cretaceous
|
Europe
|
|
|
Batrachognathus[17]
|
Rjabinin
|
1948
| Valid.
|
6 Late Jurassic.[18]
|
Asia.[18]
|
|
|
Beipiaopterus[19]
|
Lü J.-C.
|
2003
| Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Belonochasma[2]
|
Broili
|
1939
|
Misidentification.[2]
|
N/A
|
N/A
|
A non-pterosaurian gnathostome.[2]
|
|
Bennettazhia
|
Nesov
|
1991
|
Valid.
|
7 Early Cretaceous
|
N. America
|
|
|
Bogolubovia[20]
|
Nesov
A. A. Yarkov
|
1989
|
Valid.
|
8 Late Cretaceous
|
Asia
|
|
|
Boreopterus[21]
|
Lü
Qiang
|
2005
| Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Brachytrachelus
|
Giebel
|
1852
|
Preoccupied.
|
N/A
|
N/A
|
preoccupied name; now known as Scaphognathus
|
|
Brasileodactylus[22]
|
Kellner
|
1984
|
Valid.
|
7 Early Cretaceous
|
S. America
|
|
|
Cacibupteryx[23]
|
Gasparini
Fernández
de la Fuente
|
2004
| Valid.
|
6 Late Jurassic
|
Cuba
|
|
|
"Campylognathus"[24]
|
Plieninger
|
1894
|
Preoccupied.
|
N/A
|
N/A
|
preoccupied name; now known as Campylognathoides
|
|
Campylognathoides[7]
|
Strand
|
1928
| Valid.
|
5 Middle Jurassic[25]
|
Europe.[25]
|
|
|
Carniadactylus[26]
|
Dalla Vecchia
|
2009
|
Valid.
|
3 Late Triassic
|
Europe
|
Formerly Eudimorphodon rosenfeldi.[26]
|
|
Cathayopterus[27][28]
|
Wang
Zhonghe
|
2006
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Caulkicephalus[29]
|
Steel
Martill
et al.
|
2005
|
Valid.
|
7 Early Cretaceous
|
Europe
|
|
|
Caviramus [30]
|
Fröbisch
Fröbisch
|
2006
| Valid.
|
3 Upper Triassic
|
Europe
|
|
|
Cearadactylus[31]
|
Leonardi
Borgomanero
|
1985
|
Valid.
|
7 Early Cretaceous
|
S. America
|
|
|
Changchengopterus[32]
|
Lü
|
2009
| Valid.
|
6 Late Jurassic
|
Asia
|
|
|
Chaoyangopterus[33]
|
Wang
Zhou
|
2003
| Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Cimoliornis
|
Owen
|
1846
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Ornithocheirus.
|
|
Coloborhynchus[7]
|
Owen
|
1874
| Valid.
|
7 Early Cretaceous
|
Europe
N. America
S. America
|
|
|
Comodactylus[34]
|
Galton
|
1981
|
Valid.
|
6 Late Jurassic
|
N. America
|
|
|
Cretornis
|
Fritsch
|
1880
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Ornithocheirus.
|
|
Criorhynchus
|
Owen
|
1874
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Ornithocheirus.
|
|
Ctenochasma[35]
|
von Meyer
|
1852
| Valid.
|
6 Late Jurassic.[36]
|
Europe.[36]
|
|
|
Cycnorhamphus[7]
|
Seeley
|
1870
|
Valid.
|
6 Late Jurassic.[37]
|
Europe.[37]
|
|
|
"Daitingopterus"[38]
|
Maisch
Matzke
Ge Sun
|
2004
| Nomen nudum.
|
6 Late Jurassic
|
Europe
|
|
|
Darwinopterus[39]
|
Lü
Unwin
et al.
|
2009
| Valid.
|
5 Middle Jurassic
|
Asia
|
Intermediate form between rhamphorhynchoids and pterodactyloids.
|
|
Dendrorhynchoides[40]
|
Ji S.-A.
Ji Q.
Padian
|
1999
| Valid.
|
6 Late Jurassic or 7 Early Cretaceous
|
Asia
|
|
|
"Dendrorhynchus"[41]
|
Ji S.-A.
Ji Q.
|
1998
|
Preoccupied.
|
N/A
|
N/A
|
Preoccupied name; now known as Dendrorhynchoides.
|
|
Dermodactylus
|
Marsh
|
1881
| Valid.
|
6 Late Jurassic
|
N. America
|
|
|
Dimorphodon[7]
|
Owen
|
1859
|
Valid.
|
4 Early Jurassic[42]
|
Europe.[42]
|
|
|
Diopecephalus
|
Seeley
|
1871
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Pterodactylus.
|
|
Dolichorhamphus
|
Seeley
|
1880
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Rhamphocephalus.
|
|
Domeykodactylus[43]
|
Martill
Frey
et al.
|
2000
| Valid.
|
7 Early Cretaceous
|
S. America
|
|
|
Doratorhynchus[44]
|
Seeley
|
1875
|
Nomen vanum
|
6 Late Jurassic or 7 Early Cretaceous
|
Europe
|
|
|
Dorygnathus[7]
|
Wagner
|
1860
| Valid.
|
5 Middle Jurassic[45]
|
Europe.[45]
|
|
|
Dsungaripterus[7]
|
Young
|
1964
| Valid.
|
6 Late Jurassic-7 Early Cretaceous[46]
|
Africa.[46]
Asia.[46]
|
|
|
Elanodactylus[47]
|
Andres
Ji, Q.
|
2008
| Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Eoazhdarcho[48]
|
Lü
Qiang
|
2005
| Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Eopteranodon[49]
|
Lü, J.C.
Zhang
|
2005
| Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Eosipterus[7]
|
Ji
Ji
|
1997
| Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Eudimorphodon[7]
|
Zambelli
|
1973
|
Valid.
|
2 Middle Triassic[50]
|
Europe.[50]
|
|
|
Feilongus[51]
|
Wang,
Kellner,
et al.
|
2005
| Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Gallodactylus
|
Fabre
|
1974
| Jr. synonym.
| N/A
| N/A
| Jr. synonym of Cycnorhamphus.
|
|
Geosternbergia
|
Miller
|
1978
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Pteranodon.
|
|
Gegepterus[52]
|
Wang, X.
Kellner
et al.
|
2007
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Germanodactylus[7]
|
Yang
|
1964
| Valid.
|
6 Late Jurassic.[53]
|
Europe.[53]
|
|
|
Gnathosaurus[7]
|
von Meyer
|
1833
| Valid.
|
6 Late Jurassic.[54]
|
Europe.[54]
|
|
|
Haopterus[55]
|
Wang X.
Lü J
|
2001
| Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Harpactognathus[56]
|
Carpenter
Unwin
et al.
|
2003
| Valid.
|
6 Late Jurassic
|
N. America
|
|
|
Hatzegopteryx[57]
|
Buffetaut
Grigorescu, D., and
Csiki, Z.
|
2002
| Valid.
|
8 Late Cretaceous
|
Europe
|
|
|
Herbstosaurus[7]
|
Casamiquela
|
1974
| Valid.
|
6 Late Jurassic
|
S. America
|
|
|
Hongshanopterus[58]
|
Xiaolin Wang
Campos
et al.
|
2008
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Huanhepterus[59]
|
Dong
|
1982
|
Valid.
|
6 Late Jurassic
|
Asia
|
|
|
Huaxiapterus[60]
|
Lü, J., and
Yuan, C
|
2005
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Ingridia[61]
|
Unwin
Martill
|
2007
|
Jr. synonym.
|
N/A
|
N/A
|
Objective Jr. synonym of Tupandactylus.
|
|
Istiodactylus[7]
|
Howse
Milner
Martill
|
2001
|
Valid.
|
7 Early Cretaceous
|
Europe
|
|
|
Jeholopterus[62]
|
Wang, X.,
Zhou, Z.
|
2002
|
Valid.
|
5 Middle Jurassic or 7 Early Cretaceous
|
Asia
|
|
|
Jidapterus[63]
|
Dong,
Sun, &
Wu,
|
2003
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Kepodactylus[64]
|
Harris, J.D.
Carpenter
|
1996
|
Valid.
|
6 Late Jurassic
|
N. America
|
|
|
Lacusovagus[65]
|
Witton
|
2008
|
Valid.
|
7 Early Cretaceous
|
S. America
|
|
|
Laopteryx
|
Marsh
|
1881
|
Valid.
|
6 Late Jurassic
|
N. America
|
|
|
Liaoningopterus[60]
|
Wang X.-L. and
Zhou Z.-H
|
2003
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Liaoningopteryx
|
Wang X.
Zhao
|
2003
|
Lapsus calami.
|
N/A
|
N/A
|
Misspelling of Liaoningopterus.
|
|
Liaoxipterus[59]
|
Dong, Z.
Lü, J.
|
2005
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
"Lithosteornis"[66]
|
Gervais
|
1844
|
Nomen nudum.
|
N/A
|
N/A
|
Nomen nudum. Also a Jr. synonym of Ornithocheirus.
|
|
Lonchodectes[7]
|
Hooley
|
1914
|
Valid.
|
7 Early Cretaceous to 8 Late Cretaceous
|
Europe
|
|
|
Lonchognathosaurus[38]
|
Maisch, M.W.,
Matzke, A.T., and
Ge Sun
|
2004
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Longchengpterus[67]
|
Wang, L.,
Li, L
et al.
|
2006
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Ludodactylus[68]
|
Frey, E.,
Martill
Buchy, M
|
2003
| Valid.
|
7 Early Cretaceous
|
S. America
|
|
|
Macrotrachelus
|
Giebel
|
1852
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Pterodactylus.
|
|
Mesadactylus[69]
|
Jensen
Padian
|
1989
|
Valid.
|
6 Late Jurassic
|
N. America
|
|
|
Montanazhdarcho[70]
|
Padian, K.,
Horner
and de Ricqlès, A.J.
|
1993
|
Valid.
|
8 Late Cretaceous
|
N. America
|
|
|
Muzquizopteryx[71]
|
Frey, E.,
Buchy, M.-C.,
et al.
|
2006
|
Valid.
|
8 Late Cretaceous
|
N. America
|
|
|
Mythunga[72]
|
Molnar
Thulborn, R.A.
|
2008
|
Valid.
|
7 Early Cretaceous
|
Australia
|
|
|
Nemicolopterus[73]
|
Wang
Kellner
et al.
|
2008
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Nesodactylus[74]
|
Colbert
|
1969
|
Valid.
|
6 Late Jurassic
|
Cuba
|
|
|
Nesodon[75]
|
Jensen
Ostrom
|
1977
|
Lapsus calami.
|
N/A
|
N/A
|
Misspelling of Nesodactylus, also preoccupied by a toxodont.
|
|
Ningchengopterus[76]
|
Lu
|
2009
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Noripterus[7]
|
Yang
|
1973
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Normannognathus[38]
|
Buffetaut
Lepage, J.-J.
Lepage, G.
|
1998
|
Valid.
|
6 Late Jurassic
|
Europe
|
|
|
Nurhachius
|
Wang
Kellner
et al.
|
2003
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Nyctodactylus[77]
|
Marsh
|
1881
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Nyctosaurus.
|
|
Nyctosaurus[78]
|
Marsh
|
1876
| Valid.
|
8 Late Cretaceous[79]
|
N. America.[79]
S. America.[79]
|
|
|
"Odontorhynchus"
|
Stolley
|
1936
|
Preoccupied.
|
N/A
|
N/A
|
Preoccupied, but no replacement name has yet been coined.
|
|
"Oolithorhynchus"[2]
|
Whalley
|
2000
|
Nomen nudum.[2]
|
|
|
Manuscript name[2] about which almost nothing is known.
|
|
Ornithocephalus[80]
|
von Sömmering
|
1812
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Pterodactylus.
|
|
Ornithocheirus[7]
|
Seeley
|
1869
| Valid.
|
6 Late Jurassic.[81]
|
Africa.[81]
Australia.[81]
Europe.[81]
S. America.[81]
|
|
|
Ornithodesmus[2]
|
Seeley
|
1887
|
Misidentification.[2]
|
N/A
|
N/A
|
Misidentified deinonychosaur.[2]
|
|
Ornithopterus[82]
|
Fitzinger
|
1843
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Rhamphorhynchus.
|
|
Ornithostoma[7]
|
Seeley
|
1871
|
Valid.
|
8 Late Cretaceous
|
Europe
|
|
|
"Osteornis"[66]
|
Gervais
|
1844
|
Nomen nudum.
|
N/A
|
N/A
|
Nomen nudum, but appeared to be a Jr. synonym of Ornithocheirus anyway.
|
|
Pachyrhamphus[82]
|
Fitzinger
|
1843
|
Preoccupied.
|
N/A
|
N/A
|
preoccupied name; now known as Scaphognathus
|
|
Palaeornis
|
Mantell
|
1835
|
Preoccupied.
|
N/A
|
N/A
|
preoccupied name but appears to be a Jr. synonym of Ornithocheirus anyway.
|
|
Paranurognathus[83]
|
Peters
|
2005
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Anurognathus.
|
|
Parapsicephalus[84]
|
von Arthaber
|
1919
|
Valid.
|
4 Early Jurassic
|
Europe
|
|
|
Peteinosaurus[7]
|
Wild
|
1978
| Valid.
|
2 Middle Triassic[85]
|
Europe.[85]
|
|
|
"Phobetor"[2][7]
|
Bakhurina
|
1986
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Noripterus[86]
|
|
Phosphatodraco[87]
|
Pereda-Suberbiola
N. Bardet
et al.
|
2003
|
Valid.
|
8 Late Cretaceous
|
Africa
|
|
|
Plataleorhynchus[88]
|
Howse, S.C.B., and
Milner, A.R.
|
1995
|
Valid.
|
6 Late Jurassic to 7 Early Cretaceous
|
Europe
|
|
|
Preondactylus[7]
|
Wild
|
1983
|
Valid.
|
2 Middle Triassic[89]
|
Europe.[89]
|
|
|
"Pricesaurus"[90]
|
Bonaparte
Sanchez, T.M
|
1986
|
Nomen nudum.
|
7 Early Cretaceous
|
S. America
|
|
|
Procoelosaurus[91]
|
Atanassov
|
2002
|
Nomen ex dissertatione.[2]
|
|
|
|
|
Ptenodactylus[92]
|
Seeley
|
1869
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Pterodactylus.
|
|
Ptenodracon[93]
|
Lydekker
|
1888
|
Preoccupied.
|
N/A
|
N/A
|
Preoccupied name, but appears to be a Jr. synonym of Coloborhynchus anyway.
|
|
Pteranodon[7]
|
Marsh
|
1876
|
Valid.
|
8 Late Cretaceous[94]
|
N. America.[94]
|
|
|
Ptéro-dactyle
|
Cuvier
|
1809
|
|
N/A
|
N/A
|
Renamed Pterodactylus.
|
|
Pterodactylus
|
Rafinesque
|
1815
| Valid.
|
6 Late Jurassic.[95]
|
Africa.[95]
Europe.[95]
|
|
|
Pterodaustro[7]
|
Bonaparte
|
1970
| Valid.
|
7 Early Cretaceous[96]
|
S. America.[96]
|
|
|
Pteromimus[91]
|
Atanassov
|
2002
|
Nomen ex dissertatione.[2]
|
|
|
|
|
Pteromonodactylus
|
Teryaev
|
1967
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Rhamphorhynchus.
|
|
Pterorhynchus[97]
|
Czerkas
Ji, Q.
|
2002
|
Valid.
|
6 Late Jurassic
|
Asia
|
|
|
Pterotherium[98]
|
Fischer von Waldheim
|
1813
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Pterodactylus.
|
|
Puntanipterus[99]
|
Bonaparte
Sanchez, T.M
|
1975
| Valid.
|
6 Late Jurassic to 7 Early Cretaceous
|
S. America
|
|
|
Quetzalcoatlus[7]
|
Lawson
|
1975
| Valid.
|
8 Late Cretaceous[100]
|
N. America.[100]
|
|
|
Raeticodactylus[101]
|
Stecher
|
2008
|
Valid.
|
3 Upper Triassic
|
Europe
|
|
|
Rhabdopelix[2]
|
Cope
|
1870
| Misidentification.[2]
| N/A
| N/A
| At first it was thought to be a Triassic pterosaur , but is now known to be (at least in part) a kuehneosaurid.[2]
|
|
Rhamphinion[102]
|
Padian
|
1984
|
Valid.
|
4 Early Jurassic
|
N. America
|
|
|
Rhamphocephalus[7]
|
Seeley
|
1880
|
Valid.
|
midjur
|
Europe
|
|
|
Rhamphorhynchus[7]
|
von Meyer
|
1846
| Valid.
|
6 Late Jurassic.[103]
|
Africa.[103]
Europe.[103]
|
|
|
Santanadactylus[104]
|
de Buisonjé
|
1980
| Valid.
|
7 Early Cretaceous
|
S. America
|
|
|
Scaphognathus[7]
|
Wagner
|
1861
| Valid.
|
6 Late Jurassic.[105]
|
Europe.[105]
|
|
|
Sericipterus
|
|
2010
|
Valid.
|
6 Late Jurassic
|
Asia
|
|
|
Shenzhoupterus[106]
|
Lü J.
Unwin
et al.
|
2008
| Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Sinopterus[60]
|
Wang, X.
Zhou, Z
|
2003
| Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
Siroccopteryx[107]
|
Mader
Kellner
|
1999
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Coloborhynchus.
|
|
Sordes[7]
|
Sharov
|
1971
|
Valid.
|
6 Late Jurassic.[108]
|
Asia.[108]
|
|
|
Sultanuvaisia[6]
|
Nesov
|
1981
|
Misidentification.
|
8 Late Cretaceous
|
Asia.
|
Actually a fish.
|
|
Tapejara[7]
|
Kellner
|
1989
| Valid.
|
7 Early Cretaceous
|
S. America
|
|
|
Tendaguripterus[109]
|
Unwin
Heinrich
|
1999
| Valid.
|
6 Late Jurassic
|
Africa
|
|
|
Thalassodromeus[110]
|
Kellner
Campos
|
2002
|
Valid.
|
7 Early Cretaceous
|
S. America
|
|
|
"Titanopteryx"[111]
|
Arambourg
|
1959
| Preoccupied.
|
8 Late Cretaceous
|
Asia
| Preoccupied by a simuliid blackfly. It was later renamed Arambourgiania.
|
|
Tribelesodon
|
Bassani
|
1886
|
Misidentification.
|
N/A
|
N/A
|
At first it was thought to be a Triassic pterosaur but is now known to be a misinterpreted specimen of the prolacertiform Tanystropheus.
|
|
Tropeognathus
|
Wellnhofer
|
1987
|
Jr. synonym.
|
N/A
|
N/A
|
Jr. synonym of Ornithocheirus.
|
|
Tupandactylus[112]
|
Kellner
Campos
|
2007
|
Valid.
|
7 Early Cretaceous
|
S. America
|
|
|
Tupuxuara[113]
|
Kellner
Campos
|
1988
|
Valid.
|
8 Late Cretaceous
|
N. America
S. America
|
|
|
Uktenadactylus[114]
|
Rodrigues
Kellner
|
2008
|
Disputed.
|
6 Early Cretaceous
|
N. America
|
Possible junior synonym of Coloborhynchus.
|
|
Utahdactylus[115]
|
Czerkas
Mickelson
|
2002
| Disputed.
|
N/A
|
N/A
|
Probably a non-pterosaurian reptile.
|
|
Volgadraco[116]
|
Averianov
Arkhangelsky
Pervushov
|
2008
| Valid.
|
8 Late Cretaceous.
|
Asia.
|
|
|
Wukongopterus
|
|
2009
|
Valid.
|
5 Middle Jurassic
|
Asia
|
|
|
"Wyomingopteryx"[117]
|
Bakker
|
1994
| Nomen nudum.
|
|
| Nomen nudum.
|
|
Yixianopterus[118]
|
Lü, Ji S., et al.
|
2006
|
Valid.
|
7 Early Cretaceous
|
Asia
|
Appears to be valid, but little is known[119]
|
|
Zhejiangopterus[120]
|
Cai, Z.
Wei, F.
|
1994
|
Valid.
|
8 Late Cretaceous
|
Asia
|
|
|
Zhenyuanopterus[121]
|
Lü
|
2010
|
Valid.
|
7 Early Cretaceous
|
Asia
|
|
|
|
|
Although pterosaur eggs are known, some with complete embryos, no oogenera have been erected to house them. The holotype of the oospecies Oolithes sphaericus was briefly considered by Harry Govier Seeley to be pterosaurian in origin, although this attribution was dismissed before the formal erection of that oogenus.